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    3). This positively charged molecule is pushed by the membrane potential across the Hepatotoxicity Due to Mitochondrial Injury 59 Figure 6 Opposite effects of uncouplers on mitochondrial respiration and ATP formation. Uncoupling of Mitochondrial Respiration by Protonophoric Drugs Several drugs can translocate protons from the intermembrane space into the mitochondrial matrix (Fig.

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